BIOLOGY 

LIBRARY 

G 


The  North  American 

species  of 
COELAMBUS 

By  H.  C.  \FALL 


PUBLISHED  BY 

JOHN  D.  SHERMAN,  Jr. 

MOUNT  VERNON,  NEW  YORK 
1919 


Copyright,  1919 

by 
John  D.  Sherman,  Jr. 


The  North  American  Species 
of  Coelambus 

By  H.  C.  FALL 

The  members  of  the  genus  Coelambus  possess  a  facies  which  makes 
them  tolerably  easy  of  recognition  at  sight ;  they^  may  however  be 
distinguished  at  once  and  with  certainty  from  allied  genera  by  the 
raised  line  obliquely  crossing  the  epipleurae  at  base.  Our  species  are 
fairly  numerous,  the  number  known  to  me  at  present  being  nearly  twice 
that  indicated  in  the  Henshaw  List,  all  of  which  except  the  infusccv- 
tus  of  Sharp,  are  represented  in  the  material  studied.  C.  princeps, 
recently  described  from  Florida  by  Blatchley  is  not  a  Coelambus,  but 
belongs  in  all  essentials  to  Pachydrus,  a  genus  represented  by  a  small 
number  of  species  in  the  Antilles  and  Tropical  America.  Through  the 
kindness  of  its  author  I  have  been  permitted  to  examine  the  type, 
which  seems  to  me  distinct  from  any  of  the  species  given  in  Sharp's 
Monograph. 

In  the  table  of  species  which  follows,  some  emphasis  has  been  placed 
on  the  degree  of  dilatation  of  the  male  tarsi.  Since  in  several  species 
the  tarsi  of  the  male  are  very  little  wider  than  in  the  female,  or 
even  narrower  than  in  females  of  other  species,  it  is  necessary  to  have  a 
sure  means  of  recognizing  the  males.  In  the  greater  number  of  species 
the  thickening  of  the  anterior  claw  of  the  front  tarsus  of  the  male, 
though  sometimes  feeble,  is  sufficient  at  once  to  reveal  the  sex,  but 
when  this  character  fails  it  will  be  useful  to  remember  that  in  the  male 
the  basal  two  joints  of  the  front  and  middle  tarsi  are  (one  or  both) 
perceptibly  wider  than  the  third,  while  in  the  female  the  first  three 
joints  are  of  equal  width. 

Unless  otherwise  stated,  the  types  of  all  new  species  are  in  the  writer's 
collection. 

Wherever  the  order  of  species  differs  in  the  text  from  that  in  the 
table,  the  former  should  be  followed  in  cataloguing  or  in  a  cabinet 
arrangement. 

1.    Front  margined  (except  farctus),  body  broadly  ovate  and  very  con- 
vex  beneath    (except   intermedium),   color    beneath    entirely  rufous 

or    rufotestaceous 2 

Front  not  margined  (except  masculinus  and  fastidiosus) ,  form  less 
broad  and  less  convex,  body  beneath  black  (except  laccophilinus  and 
sylvanus,  size  generally  larger 3 

1 

603963 


2.  Last  ventral   foveate  in  <$,   front  tarsi   of   <$  distinctly   dilated,   size 
small  (2  to  2.4  mm.). 

Front  margined,  elytra  with  longitudinal  lateral  carina,  color 
yellow  with  black  elytral  maculation,  metasternum,  coxal  plates 
and  sides  of  abdomen  at  base  coarsely  punctate,  remainder  of 

abdomen  very  finely  and  densely  so acaroides 

Elytra    piceous    black   with    narrow    side    markings    broad- 
ening   into    a    rounded    lobe    at    middle,    reddish    brown. 

var.   marginipennis 

Front  not  margined,  elytra  not  carinate,  color  of  elytra  uni- 
formly red  brown,  head  and  thorax  paler,  body  beneath  densely 

finely    subrugosely   punctate    throughout farctus 

Last  ventral  not  foveate  in  the  male,  front  tarsi  variable,  size 
larger  (2.6  to  3.2  mm.). 

Form  broad  and.  very  convex. 

Punctures  of  elytra  unequal,  front  tarsi  of  $  dilated .liydropicus 

Punctures  of  elytra  nearly  uniform  in  size,  elytra  reddish 
brown  with   obscure   confluent   markings,    front  tarsi   of   c? 

not  dilated punctatus 

Form  less  broad  and  less  convex,  approaching  in  these  respects 

the  species  which  follow intermedius 

3.  Color  above  uniformly  reddish  brown   or   castaneous,   beneath   not 

black,  or  only  partially  so 4 

Color  above  flavo-,  rufo-  or  piceotestaceous,  or  piceous  brown,  the 
elytral  disk  often  with  broad  nubilous  markings   which   are  never 

distinctly  vittate,   body  beneath  black 5 

Color  above  fulvo-  or  rufotestaceous,  elytra  with  distinct  vittiform 
markings  (except 'in  some  forms  of  impresso punctatus.) 14 

4.  Size  a  little  larger,  form  somewhat  broader  and  more  convex,  dis- 
parity in  elytral  punctures  less  evident,  females  opaque laccophilinus 

Size  rather  smaller,  a  little  narrower  and  less  convex,  elytral  punc- 
tures evidently  unequal  toward  the  base,  females  not  opaque: sylvanus 

5.  Elytra  with  intermixed  coarser  and  finer  punctures,  which  are  more 

or  less  generally  diffused;   elytra  without  markings 6 

Elytra  nearly  uniformly  punctate,  exception  being  made  of  the 
more  or  less  evident  sutural  and  two  discal  irregular  series  of 
coarser  punctures  visible,  especially  toward  the  base,  in  many  species 7 

6.  Front  and  middle  tarsi  of  male  narrow,  not  or  scarcely  wider  than 
in  the  female. 

Size  smaller  (2.6  to  3  mm.),  form  a  little  broader,  dual  punc- 
tuation of  elytra  not  very  evident  posteriorly turbidus 

Size  larger  (3.4  to  4  mm.),  form  narrower,  dual  punctuation  dis- 
tinct throughout dispar 

Front  and  middle  tarsi  of  male  distinctly  wider  than  in  the  female, 

the  second  joint  conspicuously  widest. 

Form  a  little  broader,  color  varying  from  dull  yellow  to  brown. 

(New  England  to  Lake  Superior.)   compar 

Form  a  little  more  elongate,  color  darker.     (California.) nigrescens 

7.  Front  tarsi  of  male  at 'most  only  moderately  dilated 8 

Front  tarsi  of  male  broadly  dilated,  nearly  or  quite  as  wide  as  the 
apical  width  of  the  tibia '. 13 

2 


8.  Middle  tibiae  of  male  unmodified,  femora  similar  in  the  sexes,  the 

posterior  ones  finely  longitudinally  sub-strigose  like  the  others 9 

Middle  tibiae  of  male  sinuate  within  and  produced  inwardly  at  apex ; 
front  and  hind   femora  dissimilar  in  the  sexes,  the   latter  without 

fine  strigosity  in  either  sex pedalis 

9.  Form  narrow,  twice  as  long  as  wide,  or  even  more,  front  claw  of 
anterior  male  tarsi  not  perceptibly  modified,  color  of  elytra  nearly 
uniform  piccotestaceous  to  piceous  brown. 

Front  tarsi  of  male  narrowly  dilated,  size  smaller  (2.8  mm.) artus 

Front  tarsi  of  male  broader,  size  larger   (3.7  to  4.2  mm.) sharp! 

Form  less  elongate,  varying  from  very  distinctly  less  than,  to  nearly 
twice  as  long  as  wide,  front  claw  of  anterior  male  tarsi  perceptibly 
modified.  (Male  of  ovoideus  not  yet  known.) 10 

10.  Front  tarsi  of   male   scarcely  wider   than  the    female,   the   anterior 
claw  unmodified;  size  small    (3.1   to  3.3  mm.),    (Lake  Superior,   H. 

B.  Territory,  Manitoba. )   suturalis 

Front  tarsi  of  male  rather  narrowly  dilated,  but  evidently  wider  than 
in  the  female;  anterior  claw  of  front  tarsi  in  the  male  slightly 
thicker  and  more  abruptly  bent  at  base;  (ovoideus  provisionally 

placed,  <$  not  yet  known.) 11 

Front  tarsi  of  male  wider,  the  second  joint  conspicuously  dilated, 
anterior  claw  evidently  modified 12 

11.  Species  of  the  Lake  Superior  and  Manitoba  region. 

Color  above  nearly  uniformly  rufotestaceous,  the   sutural  edge 

of  the  elytra  blackish;  length  3.5  mm.     (Lake  Superior.) ovoideus 

Color  above  nearly  uniformly  piceo-testaceous,  the  elytral  suture 

fainty  paler  basally;  length  3.5  to  4  mm.     (Manitoba.) canadensis 

Color   of   thorax   often   in   great   part   blackish,   of    elytra   pale 
testaceous   with   a  large,   sharply   defined   piceous   discal   blotch 

behind  the  middle.     (Dakota;   Manitoba.)    sellatus 

Species  of  the  Pacific  Coast  and  Sonoran  region. 

Color  above  testaceous  with  large  elytral  cloud  extending  nearly 
to  base ;  form  slightly  narrower,  the  size  as  a  rule,  a  little  smaller 

than  the  next lutescens 

Color  nearly  as  in  the  preceding  but  with  the  elytral  cloud  less 
extended  basally  as  a  rule ;  the  elytral  markings  very  variable  in 

development,  sometimes  nearly  wanting I medialis 

Color   above   yellowish    testaceous,    elytra    with    faint    posterior 
dorsal  cloud.     Length  3.6  mm.   (Utah.)  virgo 

12.  Brownish  yellow,   the   elytra   darker   with   obscure   discal   markings 
behind  the  middle.     (Washington;  Oregon.)   obscureplagiatus 

13.  Front  tarsi  of  male  less  widely  dilated,   not  quite  as   wide  as  the 

tibial  apex collatus 

Front  tarsi  of  male  subequal  in  width  to  the  apex  of  the  tibia. 

Head  with  vertex  and  sides  of  front  more  or  less  piceous. 

Size  smaller  (3%  mm.  more  or  less.)  patruelis 

Size  larger   (5  mm.)    infuscatus 

Head  entirely  pale  except  for  a  small  transverse  occipital  spot. 
Size  larger  (over  4  mm.)   and  form  more  elongate. 

Femora  of  male  normally  feebly  sinuate  beneath  near 

the  knee,  tibiae  not  pedunculate nubilus 


Femora  of  male  strongly  sinuate  beneath  at  apex,  tibiae 

pedunculate  at  base femoratus 

Size  smaller  (less  than  4  mm.),  form  broader;  color  gen- 
erally paler  with  better  defined  elytral  clouds  behind  the 
middle fraternus 

14.  Elytra  without  impressed  lines 15 

Elytra  with  more  or  less  evident  impressed  lines  in  basal  half,  at 

least  in  the  male,  feeble  or  wanting  in  the  female 19 

15.  Punctures  of  elytra  evidently  unequal 16 

Punctures  of  elytra  denser  and  nearly  uniform  in  size 17 

16.  Last  ventral  with  an  obtuse  tumidity  each  side,  more  prominent  in 

the  male tumidiventris 

Last  ventral  not  tumid." punctilineatus 

17.  Clypeus  margined. 

Form  more  distinctly  obovate,  sides  of  thorax  not  continuous 

with  those  of  the  elytra masculinus 

Form    less    distinctly    obovate,    thorax    larger,    sides    straight, 

slightly  sinuate,  continuous  with  sides  of  elytra fastidiosus 

Clypeus  not  margined 1 8 

18.  Sides  of  thorax  feebly  divergent  posteriorly,  not  continuous  with 
sides   of  elytra:    elytral  vittae   narrow,   much  abbreviated  in   front. 

semivittatus 

Sides  of  thorax  normally  divergent,  nearly  continuous  with  sides  of 
elytra. 

Size  smaller  (3.6  to  4  mm.),  claws  of  front  tarsi   (<£)   unusually 

elongate,  not  very  unequal  in  length  or  thickness. hudsonicus 

Size  larger   (5  to  5.8  mm.) 

Form   broader,    surface   beneath    alutaceous    in   both    sexes, 

claws  of  front  tarsi   (<£)  very  unequal unguicularis 

Form  more  elongate,  surface  beneath  not  alutaceous,  claws 

of  front  tarsi    (c?)    stout,  rather  short,  equal oregonus 

19.  Size  larger   (4.5  to  5.5  mm.),  color  varying  from  pale  yellow,  with 
vittate  elytra  to  nearly  uniformly  piceous;   punctuation  very  coarse 

with  intermixed  fine  punctures,  less  obvious  in  the  female 

impressopunctatus 

C.  acaroides  Lee. 

Rotundate  oval,  rufotestaceous,  body  beneath  often  darker,  brownish  or  even 
piceous  with  the  prothorax  and  abdominal  apex  pale ;  elytra  with  the  suture 
narrowly,  a  broader  more  or  less  abbreviated  subsutural  stripe  (often  inter- 
rupted), and  two  external  discal  spots,  blackish;  the  markings  are,  however, 
very  variable  through  confluence  or  suffusion.  Antennae  and  legs  pale,  the 
tarsi  sometimes  dusky.  Head  and  thorax  finely,  not  very  closely  punctate ;  elytra 
more  coarsely  and  closely  and  very  evenly  so.  Coxal  plates  very  coarsely 
closely  punctate,  abdominal  segments  very  minutely  densely  punctulate  and  dull, 
the  basal  segments  at  sides  with  scattered  shallow  coarser  punctures.  Clypeus 
very  finely  margined ;  elytra  with  a  fine  acute  submarginal  carina  extending 
forward  from  the  middle  but  not  attaining  the  base.  Length  2.1 'to  2.4  mm.; 
width  1.3  to  1.5  mm. 

Var.  marginipennis  Blatch. 

Similar  in  all  essentials  to  the  "typical  form,  but  with  an  extreme  suffusion  of 


the  elytral  spots,  leaving  only  a  pale  lateral  margin  which  broadens  into  a 
rounded  lobe  at  the  middle.  Blatchley  in  his  description  says  there  is  no  elytral 
carina,  it  is  however,  quite  as  distinct  in  a  typical  example  before  me  as  in  the 
usual  acaroides  form. 

The  types  of  acaroides  were  from  Illinois  and  Kentucky.  Specimens 
before  me  are  from  Illinois ;  Miss ;  Fedor,  Texas ;  Winnfield  La. 
(Pilate.)  It  is  reported  from  Cincinnati,  Ohio,  by  Dury,  and  is  on  the 
Iowa  List(Wikham.)  The  variety  occurs  in  Florida  (Sarasota  and 
Dunedin — Blatchley. ) 

The  front  and  middle  tarsi  of  the  male  are  distinctly  dilated,  and 
there  is  in  this  sex  a  small  discal  excavation  of  the  last  ventral  from 
the  anterior  margin  of  which  project  backward  two  approximate 
porrect  slender  spines.  This  remarkable  structure  exists  in  the  follow- 
ing species,  but  so  far  as  I  know  has  no  parallel  elsewhere  in  the 
family,  and  seems  to  have  been  entirely  overlooked. 

C.  farctus  Lee. 

Rotundate  oval,  feebly  shining  above,  opaque  beneath,  general  color  reddish 
brown,  the  elytra  darker  brown  with  the  side  margins  suffusedly  paler,  outer 
joints  of  antennae,  tarsi  and  frequently  the  basal  half  of  the  abdomen  more 
or  less  infuscate.  Head  not  margined  in  front,  finely  densely  punctate. 
Prothorax  similarly  punctate,  a  few  interspersed  coarser  punctures  along  the 
base.  Elytra  finely,  closely  punctate,  with  numerous  intermixed  coarser  ocellate 
punctures  in  the  basal  half,  these  becoming  less  numerous  or  nearly  disappearing 
posteriorly.  Body  beneath  very  densely  punctate,  the  ventral  segments  very 
minutely  so,  the  metasternum  and  coxal  plates  more  coarsely  so.  Length  2.2 
to  2.3  mm.;  width  1.35  to  1.4  mm. 

Le  Conte's  type  was  from  Massachusetts,  and  all  the  specimens  at 
hand  are  from  Tyngsboro,  Mass.,  where  I  have  taken  it  somewhat 
sparingly  in  a  small  woodland  pond.  Crotch  recorded  it  from  New 
Jersey  also  and  recent  collectors  report  its  occurrence  at  Lakehurst 
in  that  state. 

The  four  anterior  tarsi  are  quite  distinctly  dilated  in  the  male, 
evidently  narrower  in  the  female.  The  last  ventral  of  the  male  shows 
the  same  median  excavation  and  porrect  spinules  as  in  acaroides,  but 
the  excavation  is  here  rather  larger  and  the  spinules  less  approximate. 

C.  punctatus  Say. 

Broadly  rotundate  ovate ;  very  convex,  especially  beneath,  head  thorax  and  body 
beneath  rufotestaceous,  the  elytra  usually  duller  brownish  yellow,  the  disk  with 
obscure  fuscous  markings  which  are  sometimes  feebly  defined,  but  usually 
broadly  suffused;  base  and  apex  of  thorax  blackish;  antennae  and  legs  entirely 
rufotestaceous.  Punctuation  above  strong,  close  and  nearly  uniform,  beneath 
coarse  and  close.  Tarsi  narrow,  not  appreciably  broader  in  the  male.  Length 
2.8  to  3  mm.;  width  1.7  to  1.9  mm. 

A  common  and  well  known  species  widely  dispersed  in  Canada  and 
the  northern  half  of  the  L^nited  States  from  New  England  to  the 


Pacific  coast.  Specimens  are  known  to  me  or  reliably  reported  from 
Maine,  (York  Beach;)  Massachusetts  (Marion,  Tyngsboro;)  New 
York  (Adirondack  Mountains;  Buffalo;)  New  Jersey  ("all  sections  of 
state" — Smith's  List;)  Southwestern  Pennsylvania  (Hamilton  List;) 
Ohio;  N.  Illinois;  Wisconsin;  Central  and  Northern  Indiana 
(Blatchley;)  Ottawa,  Canada;  Cochrane,  N.  Ontario  (Notman;)  Lake 
Superior;  Winnipeg,  Manitoba  (Wallis,)  Kansas;  "Northwest  Terri- 
tory" (Say's  description;)  British  Columbia  (Peachland — Wallis;) 
Seattle,  Washington  (O.  B.  Johnson;)  Carlin,  Nevada. 

This  species  has  been  held  by  most  writers  to  be  at  most  only  a 
variety  of  the  European  inaequalis.  This  view  however  is  not  shared 
by  Sharp  who  says  of  punctatus  "rather  different  in  outline  being  broader 
in  the  middle  and  so  less  parallel  in  form,  with  the  punctuation  of  the 
elytra  rather  dense,  and  the  upper  surface  without  distinct  contrasts 
of  color."  The  difference  in  form  stated  by  Sharp  seems  to  have  some 
slight  basis  in  fact,  though  rather  difficult  of  appreciation  by  the 
unimaginative  student.  Personally  I  have  seen  no  American  examples 
that  in  brightness  of  makings  could  be  considered  as  typical  Inaequalis, 
Crotch  however  mentions  "two  specimens  from  Illinois  marked  precisely 
as  in  the  European  species." 

C.  hydropicus  Lee. 

Broadly  ovate,  convex,  nearly  similar  in  form  to  punctatus  but  a  little  more 
elongate,  flavo-or  rufotestaceous,  outer  joints  of  the  antennae  and  front  and 
hind  margins  of  thorax  more  or  less  infuscate;  elytra  dull  yellow,  disk  with 
broadly  confluent  fuscous  shades,  which  in  the  more  definitely  marked  examples 
leave  the  sides  and  transverse  basal  and  postmedian  series  of  spots  or  dashes, 
yellow.  Punctuation  above  and  beneath  still  denser  than  in  punctatus,  and  on 
the  elytra  distinctly  dual  in  character,  especially  baso-medially ;  not  very  ob- 
viously so  in  punctatus.  Front  and  middle  tarsi  very  distinctly  dilated  in  the 
male,  the  claws  not  sexually  modified.  Length  2.8  to  3.2  mm.;  width  1.7  to  1.8 
mm. 

California — Middle  and  Southern.  Le  Conte's  type  is  from  San 
Diego.  Specimens  before  me  are  from  Pomona,  Pasadena,  Santa 
Margarita  (San  Luis  Obispo  Co. — Martin)  Guerneville,  Sonoma  Co. 
(Blaisdell,)  and  Vine  Hill,  Contra  Costa  Co.  (Blaisdell.) 

The  general  similarity  to  the  well  known  punctatus,  combined  with 
the  dilated  male  tarsi  render  this  species  easily  recognizable.  It 
is  thus  far  known  to  me  only  from  California  and  I  have  as  yet  seen 
no  specimens  of  punctatus  from  this  state,  though  its  proximity  both  to 
the  north  and  east  make  its  ultimate  discovery  there  not  unlikely. 

C.  intermedius  sp.  nov. 

Oval,  moderately  elongate,  yellowish  testaceous,  head  (more  or  less)  and  front 
and  hind  margins  of  thorax  infuscate;  elytra  with  broad  suffused  fuscous 
clouds  which  leave  only  the  side  margins  and  some  indistinct  subbasal  and 

6 


postmedian  spots  paler.  Integuments  polished  throughout.  Antennae  scarcely 
infuscate  apically.  Head  margined  in  front.  Punctuation  of  head  and  thorax 
fine,  not  very*  close;  elytra  more  coarsely  and  closely  punctate.  Body  beneath 
rather  strongly  and  not  very  closely  punctate,  the  punctures  coarser  on  the 
coxal  plates  as  usual.  Length  2.6  mm.;  width  1.4  mm. 

The  type  was  collected  in  Alameda  Co.,  California  by  Koebele.  The 
specimen  is,  I  think,  a  male,  though  the  tarsi  are  only  very  moder- 
ately dilated.  A  second  specimen  is  from  Dalles,  Oregon. 

This  litle  species  is  by  the  margined  front  allied  to  those  which  pre- 
cede, but  is  distinctly  less  convex  both  above  and  beneath,  and  has 
the  moderately  elongate  oval  form  of  most  of  the  species  which  follow. 

C.  laccophilinus  Lee. 

Ovate,  pointed  behind,  moderately  convex,  rufocastaneous,  coxal  plates  and 
base  of  abdomen  sometimes  infuscate.  Integuments  sparsely  rather  finely 
punctate,  strongly  shining  in  the  male,  alutaceous  and  opaque  and  more  finely 
punctate  in  the  female.  Length  2.8  to  3.1  mm.;  width  1.65  to  1.75  mm. 

Described  from  Detroit,  Michigan ;  known  to  me  also  from  Tyngs- 
boro,  Mass.,  Peekskill,  N.  Y.  (Sherman),  and  Indiana  (Blatchley.) 
It  is  also  recorded  by  Smith  in  the  New  Jersey  List. 

This  species  is  similar  in  many  respects  to  the  next,  but  is  a  little 
larger,  broader  and  more  convex,  and  with  less  inequality  in  size  of 
punctures.  The  front  and  middle  tarsi  are  very  distinctly  wider  in 
the  males  than  in  the  females,  the  disparity  being  much  less  marked  in 
sylvaiius. 

C.  sylvanus  Fall. 

Form  rather  narrowly  ovate,  widest  before  the  middle,  moderately  convex, 
glabrous,  both  sexes  shining;  color  castancous,  the  antennae  legs  and  under 
side  of  head  and  prothorax  brighter  rufous.  Head  and  prothorax  finely 
sparsely  punctulate,  the  latter  more  strongly  and  closely  so  along  the  front  and 
rear  margins.  Elytra  widest  a  little  behind  the  base,  sides  continuous  in  out- 
line with  the  prothorax;  punctuation  not  very  close,  consisting  of  intermixed 
moderately  coarse  and  much  finer  punctures.  Metasternum,  coxal  plates  and 
sides  of  the  basal  ventral  segments  sparsely  punctate,  the  punctures  gradually 
diminishing  in  size  from  front  to  rear.  Front  and  middle  tarsi  of  male  only 
moderately  dilated,  a  little  narrower  in  the  female ;  claws  small  and  slender, 
unmodified  in  the  male.  Length  2.5  to  2.7  mm.:  width  1.35  to  1.45  mm. 

New  York  (Peekskill— Sherman.) 

Nearly  allied  to  laccophilinus  but  smaller,  narrower  and  less  convex 
(not  "more  convex"  as  was  stated  by  mistake  in  the  original  descrip- 
tion), and  with  less  dilated  male  tarsi,  the  sexual  difference  in  this 
respect  not  very  marked. 

C.  turbidus  Lee. 

Regularly  oval,  moderately  convex,  widest  at  middle,  above  dull  yellowish  to 
brownish  or  piceotestaceous,  without  markings ;  the  head,  thorax  anteriorly,  and 
side  margins  of  elytra  a  little  brighter  in  color;  integuments  polished  in  both 


sexes;  antennae  with  outer  joints  lightly  infuscate.  Head  and  thorax  finely 
sparsely  punctate,  a  few  coarser  punctures  along  the  base  of  the  latter.  Elytra 
with  a  dual  system  of  finer  and  coarser  punctures,  which  is,  however,  not  very 
strongly  marked,  the  disparity  becoming  quite  feeble  posteriorly.  Body  beneath 
black,  metasternum,  coxal  plates  and  sides  of  abdomen  at  base  coarsely,  not 
densely  punctate,  venter  elsewhere  finely  sparsely  punctate;  under  side  of 
prothorax,  epipleurae  and  legs  pale.  Length  2.7  to  3  mm.;  width  1.5  to  1.75  mm. 

Described  by  Le  Conte  from  Massachusetts.  Crotch  says  more 
specifically,  "Boston."  It  occurs  rarely  at  Tyngsboro,  Mass.,  and  other 
Massachusetts  specimens  in  my  collection  are  labeled  Westport,  Berke- 
ley and  Fall  River  (N.  S.  Easton).  I  have  also  before  me  examples 
from  Vermont,  Montreal,  Can.;  Winnipeg,  Man.  (J.  B.  Wallis),  and 
Ag.  Coll.,  Miss.  (H.  E.  Weed.)  It  is  reported  from  Staten  Island 
(Leng),  and  from  Indiana  (Blatchley.) 

C.  dispar  Lee.    (=dissimilis  G.  &  H.) 

Moderately  elongate,  oval,  dull  yellow  to  brownish  testaceous,  head  and  thorax 
commonly  brighter  rufotestaceous,  the  typical  dorsal  discal  clouds  feebly  de- 
fined on  the  elytra  in  the  light  colored  examples,  but  scarcely  at  all  evident 
.  in  the  darker  specimens ;  trunk  black  beneath ;  integuments  polished  throughout 
in  both  sexes;  outer  joints  of  antennae  lightly  infuscate,  with  their  bases  paler. 
Head  and  prothorax  finely  sparsely  punctate,  the  latter  with  a  few  coarser 
punctures  near  the  base.  Elytra  with  conspicuously  intermixed  fine  and  much 
coarser  punctures  over  the  greater  portion  of  the  disk,  the  punctures  closer 
and  more  nearly  equal  in  size  at  apex.  Metasternum,  coxal  plates  and  the 
sides  of  the  basal  ventral  segments  coarsely  punctate.  Length  3.4  to  4  mm.; 
width  1.85  to  2.15  mm. 

This  species  is  widely  dispersed  in  the  northern  United  States  and 
Canada.  Described  from  Lake  Superior.  Specimens  before  me  are 
from  Esopus,  N.  Y. ;  Phila  Neck,  Pa. ;  South  Haven,  Mich. ;  Evanston, 
Ind. ;  N.  111. ;  Glencoe,  111. ;  Rochester,  Minn. ;  St.  Louis,  Mo. ;  Winni- 
peg, Man. ;  Kamloops,  B.  C.  It  is  recorded  also  in  the  New  Jersey, 
District  of  Columbia  and  Iowa  Lists. 

In  some  30  or  40  specimens  examined  there  are  no  obvious  sexual 
differences,  the  tarsi  being  narrow  and  of  the  female  type  in  all, 
though  there  can  be  little  doubt  that  both  sexes  are  present. 

C.  compar  sp.  nov. 

Elongate  oval,  very  similar  in  form  and  size  to  the  preceding  species,  and  like 
that,  varying  in  color  from  dull  yellow  or  rufotestaceous  to  fuscotestaceous  or 
brownish,  the  elytra,  however,  without  trace  of  nubilous  markings  which  are 
sometimes  somewhat  defined  in  dispar,  antennae  infuscate  from  about  the 
middle.  Elytra  with  intermixed  punctuation,  the  coarser  punctures,  however, 
are  smaller  than  in  dispar,  and  the  disparity  is  therefore  less  marked  and  is  not 
very  evident  except  in  the  baso-sutural  region.  The  males  are  polished,  the 
females  may  be  either  shining  or  dull,  the  latter  as  usual  more  finely  punctate. 
In  the  male  the  front  and  middle  tarsi  are  distinctly  broader,  the  second  joint 
rather  conspicuously  widest  and  triangular,  the  third  joint  about  as  long  as 
wide  and  very  slightly  wider  than  the  basal  joint,  the  anterior  claw  of  the  front 


tarsus  thicker  and  more  strongly  bent.  In  the  female  the  tarsi  are  narrow  with 
joints  1-3  equal  in  width.  Length  3.6  to  4  mm.;  width  1.95  to  2.15  mm. 

The  type'of  this  species  is  a  male  taken  by  the  writer  at  Tyngsboro, 
Mass.,  7-21-06.  Similar  examples  from  Bennington  Co.,  Vt,  are  in 
the  Roberts  and  Sherman  Collections.  Mr.  Sherman  has  also  taken  it 
at  Starr  Lake,  4,890  feet,  and  at  5,000  feet  on  Mt.  Washington,  N.  H., 
and  at  Marquette,  Mich.  In  the  series  of  dispar  in  the  Le  Conte  Col- 
lection are  two  examples  of  this  species,  one  from  "N.  H.",  the  other 
bearing  the  gray  color  disk  which  signifies  Lake  Superior. 

C.  nigrescens  sp.  nov. 

Two  examples  (c?$)from  California,  are  very  close  to  the  preceding,  but  seem 
to  be  distinct.  They  are  of  a  slightly  more  elongate  elliptico-oval  form  (more 
obtuse  at  apex)  and  of  very  dark  color;  the  head  and  prothorax  black,  elytra 
piceous  brown,  femora  broadly  infuscate  at  middle,  the  tarsi  blackish.  The 
sexual  characters  of  the  male  tarsi  are  as  in  compar,  but  the  large  second  joint 
is  even  slightly  more  developed.  Length  3.5  to  4  mm.;  width  1.8  to  1.95  mm. 
The  smaller  measurements  are  those  of  the  female. 

Mono  Co.,  California   (Blaisdell),  type  c?. 

C.  artus  sp.  nov. 

Narrowly  subelliptical ;  fuscotestaceous  above,  the  middle  of  the  front,  disk  of 
prothorax  and  elytral  margins  narrowly,  somewhat  paler;  tarsi  and  outer 
joints  of  antennae  dusky;  body  black  beneath,  integuments  polished  throughout. 
Head  and  thorax  finely  sparsely  punctate,  elytra  more  closely  and  less  finely 
punctured,  the  usual  three  series  of  coarser  punctures  very  irregular,  visible 
in  basal  half.  Body  beneath  rather  coarsely,  not  densely  punctate  at  sides. 
Length  2.9  mm.;  width  1.35  mm. 

Mono  Co.,  California  (Blaisdell.) 

The  type  and  only  specimen  at  hand  is  a  male,  but  the  tarsi  are  quite 
narrow,  the  sex  being  revealed  by  the  visibly  modified  front  claw.  The 
small  size,  exceptionally  narrow  form  and  narrow^  male  tarsi  are  the 
diagnostic  characters  to  be  relied  upon  in  this  species. 

C.  sharp!  VdBr. 

Oval,  strongly  elongate;  head  black  with  pale  spot  on  the  vertex;  thorax  rufo- 
testaceous  with  front  and  rear  margins  infuscate,  a  small  discal  spot  behind  the 
middle,  usually  confluent  with  the  dark  margin;  elytra  brownish  fuscous  or 
fuscotestaceous,  usually  nearly  uniformly  so,  sometimes,  however,  with  the 
margins  paler,  showing  vaguely  the  outlines  of  the  broadly  suffused  discal 
markings;  beneath  black,  legs  and  antennae  %pale,  the  latter  with  the  outer  joints 
slightly  dusky.  Integuments  polished,  both  above  and  beneath.  Punctuation 
fine,  not  close,  nearly  uniformly  distributed  on  head  and  thorax,  a  little  coarser 
and  denser  on  the  elytra,  the  punctures  gradually  closer  apically;  discal  lines 
of  coarser  punctures  distinct  and  quite  regular.  Metasternum  and  sides  of 
abdomen  at  base  coarsely  closely  punctate,  coxal  plates  somewhat  less  coarsely 
so.  Length  3.7  to  4.3  mm.;  width  1.8  to  2.1  mm. 

The  type  of  this  species  is  the  lutesccns  of  Sharp's  Monograph, 
which  was  doubtfully  identified  by  him  as  Le  Conte's  species,  the  de- 

9 


termination  based  largely  no  doubt  on  the  locality  (San  Francisco), 
which  is  the  same  as  for  Le  Conte's  type.  Although  very  briefly 
characterized,  the  size,  color,  and  narrow  form,  combined  with  the 
locality,  leave  no  doubt  as  to  what  Sharp  had  in  hand.  This  species 
is  thus  far  known  only  from  Central  and  Northern  California,  the 
localities  represented  in  the  material  at  hand  being — San  Francisco, 
Vine  Hill,  Contra  Costa  Co.  (Blaisdell)  ;  Santa  Cruz,  Napa,  Santa 
Rosa,  Humboldt  Co.  The  front  and  middle  tarsi  of  the  male  are 
distinctly  dilated,  the  first  and  second  joints  subequal  in  width,  the 
third  a  little  narrower ;  claws  of  front  tarsi  small,  of  the  same  form 
and  length,  the  anterior  one  barely  visibly  thicker. 

C.  suturalis  Lee. 

Oval,  nearly  equally  narrowed  before  and  behind ;  dull  yellowish  testaceous 
above;  black  beneath;  head  infuscate  near  the  eyes,  leaving  a  triangular  frontal 
area  pale ;  thorax  witth  fuscous  median  spot  confluent  with  the  dark  basal 
margin;  disk  of  elytra  broadly  infuscate,  the  markings  in  the  typical  form 
completely  suffused,  leaving  only  the  margins  and  a  narrow  sutural  line, 
paler.  In  many  specimens  the  elytral  markings  are,  however,  fairly  well 
defined.  Punctuation  rather  dense,  coarser  than  in  ovoidcus,  the  irregular 
series  of  coarser  elytral  punctures  not  very  distinct.  Beneath  coarsely,  rather 
densely  punctate.  Legs  either  uniformly  pale  or  with  the  femora  somewhat 
clouded.  Length  3.1  to  3.3  mm.;  wridth  1.7  to  1.8  mm. 

The  type  series  consists  of  three  specimens  taken  at  Lake  Superior, 
the  precise  locality  not  named.  Le  Conte  regarded  all  these  as  fe- 
males, but  a  careful  examination  satisfies  me  that  two  of  them  are 
males.  In  this  sex  the  tarsi  are  very  slightly  wider,  the  second  joint 
sensibly  wider  than  the  third,  while  in  the  female  joints  1-3  are  of 
nearly  equal  width.  The  front  claw  of  the  fore  tarsus  is  not  at  all 
modified  in  the  male.  There  is  a  fourth  specimen  from  "H.  B.  T." 
in  the  Le  Conte  Collection,  apparently  differing  from  the  original  series 
only  in  the  better  defined  elytral  markings.  I  have  seen  many  examples 
like  this  last  taken  by  Mr.  J.  B.  Wallis  in  Manitoba.  (Miami,  Le  Pas, 
Piquitenay  River  and  Mile  214  H.  B.  Ry.)  and  others  taken  by  Mr. 
Howard  Notman  at  Cochrane  in  N.  Ontario. 

C.  ovoideus  Lee. 

Oval,  about  equally  narrowed  before  and  behind,  uniformly  rufotestaceous  above, 
head  infuscate  with  pale  vertex  spot,  thorax  with  basal  margin  darker,  elytra 
with  the  sutural  edge  blackish,  body  beneath  black.  Punctuation  rather  finer 
and  less  close  than  in  the  allied  species,  the  series  of  coarser  punctures  on  the 
elytra  evident  though  not  very  conspicuous.  Length  3.5  mm.;  width  1.8  to 
1.9  mm. 

The  type  locality  is  Eagle  Harbor,  Lake  Superior.  There  are  only  two 
examples  in  the  Le  Conte  Collection,  both  of  which  judging  from  the 
tarsi  seem  to  be  females.  Le  Conte  in  his  description  speaks  of  the 
male  as  being  a  little  more  shining  than  the  female,  but  it  may  be  that  he 

10 


was  in  error  in  his  supposed  separation  of  the  sexes.  The  species  is  ap- 
parently rare  and  I  have  seen  no  other  specimens  than  the  types.  In  the 
absence  of  Tnales,  the  tabular  position  is  tentative  only. 

C.  canadensis  sp.  nov. 

Oval,  moderately  elongate,  color  above  a  nearly  uniform  piceo-  or  fuscotestace- 
ous,  fading  into  a  dingy  yellow  at  the  extreme  margins  of  the  elytra,  the 
prothorax  commonly  entirely  dark,  but  sometimes  with  the  middle  narrowly 
paler  toward  the  sides;  head  with  or  without  a  pale  vertex  spot;  body  beneath 
black;  legs  and  antennae  pale,  the  latter  infuscate  apically.  Punctuation  through- 
out typical  of  the  group,  a  little  sparser  and  finer  than  in  suturalis  or  medialis, 
nearly  the  same  as  in  patrucHs;  the  irregular  series  of  coarser  punctures  on 
the  elytra  present  but  not  conspicuous.  Length  3.5  to  4  mm.;  width  1.95  to 
2.1  mm. 

Described  from  a  series  of  23  examples  taken  at  Winnipeg,  Man., 
by  Mr.  J.  B.  Wallis.  The  fore  and  middle  tarsi  of  the  male  are  only 
moderately  dilated,  though  evidently  wider  than  in  the  female,  the 
anterior  claw  of  the  fore  tarsus  a  little  thicker  and  slightly  shorter. 
Females  are  typically  shining  like  the  males,  but  two  examples  of  this 
sex  bearing  same  date  as  shining  specimens  of  both  sexes,  are  opaque. 
These,  however,  are  not  distinguishable  from  opaque  females  of 
patruelis  and  may  belong  to  that  species,  which  is  commonly  taken  by 
Mr.  Wallis  in  the  same  locality.  Canadensis  is  very  similar  in  size  and 
form  to  patruelis,  but  is,  as  a  rule,  of  darker  and  more  uniform  color, 
and  with  distinctly  narrower  male  tarsi. 

C.  sellatus  Lee. 

Oval,  nearly  equally  narrowed  before  and  behind ;  color  above  pale  testaceous, 
beneath  black.  Head  blackish  each  side,  a  pale  vertex  spot,  and  the  clypcus  an- 
teriorly diffusely  paler.  Thorax  with  front  margin  more  narrowly,  base  broadly 
infuscate,  also  a  diffuse  discal  spot  confluent  with  the  dark  margins.  Elytra 
with  a  large  fuscous  blotch  occupying  the  posterior  portion  of  the  disk,  and  in 
front  of  this  a  small  elongate  spot  nearer  the  side  margin  than  the  suture. 
Punctuation  moderate,  coarser  beneath  as  usual,  the  irregular  series  of  larger 
punctures  on  the  elytra  variable  in  development.  Length  3  to  3.25  mm. ;  width 
1.65  to  1.75  mm. 

Described  from  Dakota.  Specimens  at  hand  are  from  Winnipeg 
and  Stony  Mountain,  Manitoba  (Wallis.)  This  species  may,  I  think, 
always  be  known  by  the  rather  sharply  defined  posterior  elytral  blotch, 
contrasting  strongly  with  the  pale  ground  color  of  the  elytra.  It  is 
rather  strongly  narrowed  in  front,  nearly  as  in  ovoid  ens.  Siituralis 
is  less  narrowed  and  consequently  more  obtuse  in  front,  with  more  ex- 
tended elytral  cloud  and  unmodified  male  anterior  tarsal  claw.  In 
sellatus  the  male  tarsi  are  rather  narrow,  but  evidently  wider  than  in 
the  female,  and  the  anterior  claw  is  visibly  stouter. 

C.  lutescens  Lee. 

Typically  a  trifle  smaller  and  narrower  than  medialis,  with  the  elytral  disk  more 

11 


completely  infuscate,  the  markings  reaching  nearly  or  quite  to  the  base;  all 
else  substantially  as  in  medialis.  Some  examples  are  so  nearly  intermediate  be- 
tween typical  forms  of  these  two  so-called  species  that  their  disposition  is  little 
more  than  a  matter  of  guess  work.  I  have  observed  in  the  numerous  specimens 
studied  that  in  lutescens  the  dark  color  at  the  sides  of  the  head  in  the  great 
majority  of  specimens  becomes  suffused  across  the  front  so  as  to  leave  a 
more  or  less  isolated  pale  vertex  spot,  while  in  medialis,  as  a  rule,  the  pale 
frontal  triangle  reaches  from  the  clypeal  margin  to  the  vertex.  Length  3.2  to 
3.5  mm.;  width  1.65  to  1.8  mm. 

Although  the  figures  giving  the  extremes  in  length  are  the  same  for 
this  species  as  for  medialis,  here  the  average  length  is  nearer  the  lower 
limit,  while  in  medialis  the  reverse  is  true. 

Le  Conte's  type  of  lutescens  was  from  San  Francisco.  Specimens 
before  me  are  from  Sacramento ;  Athlone,  Merced  Co. ;  and  Kernville, 
all  in  California. 

C.  medialis  Lee. 

Oval,  color  varying  from  albotestaceous  to  brownish  yellow  above,  black  be- 
neath; head  more  or  less  infuscate  at  sides;  prothorax  with  a  small  median 
fuscous  spot,  rarely  entirely  lacking,  front  and  rear  margins  slightly  or  scarcely 
infuscate,  more  rarely  broadly  so;  elytra  with  broad  suffused  fuscous  markings, 
which  usually  fail  to  attain  the  base,  and  are  not  infrequently  so  faint  as  to 
be  only  traceable  in  certain  lights.  Punctuation  moderately  close,  becoming 
gradually  a  little  coarser  and  denser  posteriorly  on  the  elytra,  the  sutural  and 
first  discal  series  of  coarser  punctures  quite  irregular  and  conspicuous,  the 
second  (intrahumeral)  usually  less  marked;  beneath  coarsely  punctate.  Length 
3.2  to  3.5  mm.;  width  1.75  to  2  mm. 

A  common  species,  ranging  from  Middle  and  Southern  California 
to  Western  Texas. 

The  type  is  from  San  Diego,  Cal.  Specimens  before  me  are  from 
San  Juan,  Orange  Co. ;  Pomona ;  Elsinore  Lake ;  Santa  Margarita ; 
Santa  Cruz ;  Vine  Hill,  Contra  Costa  Co. ;  San  Francisco ;  Owen's 
Lake ;  California,  Williams  and  Cosnino  Canon,  Arizona ;  Carlin, 
Nevada;  El  Paso  Texas. 

The  tarsi  are  but  little  wider  in  the  male  than  in  the  female ;  the 
anterior  claw  of  front  tarsus  (c?)  evidently  a  little  thicker  and  more 
suddenly  bent. 

"Specimens  from  Arizona  and  Texas  are  nearly  uniformly  pale  in 
color,  the  elytral  markings  faint  or  nearly  wanting ;  these  also  seem 
as  a  whole  slightly  more  finely  punctate,  but  there  are  perfect  inter- 
mediates between  them  and  the  typical  California  coast  form  with 
heavily  clouded  elytra.  These  latter  are  very  near  lutescens,  and  in 
many  cases  separation  is  purely  an  arbitrary  matter. 

C.  virgo  sp.  nov. 

Moderately  elongate  oval,  both  sexes  shining ;  head  and  thorax  brownish,  the 
former  with  pale  vertex  spot,  the  latter  with  sides  paler;  elytra  luteous,  the 

12 


dorsal  cloud  broad  but  faint,  abbreviated  in  front.  Punctuation  above  moder- 
ately close,  the  series  of  coarser  elytral  punctures  visible  but  not  conspicuous. 
Beneath  black,  coarsely  but  not  very  closely  punctate.  Front  tarsi  moderately 
dilated,  the  first  and  second  joints  subequal  in  width,  anterior  claw  slightly 
thicker,  more  strongly  bent  and  a  little  shorter  than  the  posterior  one.  Length 
3.6  to  3.7  mm.;  width  1.9  to  1.95  mm. 

Described  from  ic?  (type)  and  3?'s  from  the  Virgin  River,  Utah, 
the  type  series  from  the  Roberts  Collection  kindly  loaned  by  the 
American  Museum  of  Natural  History,  to  which  the  type  is  returned. 
A  female  paratype  is  retained  in  my  own  collection. 

This  species  is  of  same  size  and  general  appearance  as  the  paler 
forms  of  medialis,  but  is  of  slightly  narrower  form,  the  sides  being 
less  strongly  rounded,  the  punctuation  beneath  not  so  coarse  and  close, 
and  the  male  tarsi  distinctly  wider.  It  is  really  much  more  closely 
related  to  canadensis,  but  the  clearer  yellow  color,  with  evident  though 
faintly  defined  posterior  elytral  cloud  seems  at  present  distinctive. 

C,  obscureplagiatus  sp.  nov. 

Elongate  oval,  about  equally  narrowed  in  front  and  rear,  brownish  ferruginous 
or  brownish  yellow,  the  head  a  little  paler  in  front;  thorax  and  elytra  variable, 
the  former  nearly  uniformly  brown  or  piceous  brown,  the  latter  with  the  disk 
usually  dark  brownish  with  obscure  piceous  markings  of  the  niedialis  type,  the 
base  and  sides  of  the  elytra  gradually  somewhat  paler.  Surface  polished  both 
above  and  beneath  in  the  male,  the  females  either  shining  like  the  males,  or 
alutaceous  and  opaque.  Head  and  thorax  finely  typically  punctate,  elytra  more 
coarsely  quite  densely  and  nearly  uniformly  so,  the  series  of  coarser  punctures 
represented  only  by  a  lightly  impressed  line  at  the  middle  of  the  disk  in  basal 
half,  scarcely  detectable  in  some  examples.  Body  beneath  black,  coarsely 
closely  punctate.  Front  and  middle  tarsi  of  male  moderately  widely  dilated, 
the  second  joint  conspicuously  widest,  the  anterior  claw  a  little  stouter  and  more 
abruptly  curved  basally.  Length  3.8  to  3.9  mm.;  width  1.95  mm. 

Described  from  a  series  of  2c?'s  and  5?'s  (3  shining,  2  opaque) 
from  Port  Townsend,  Wash.  (A.  Seaton),  and  Portland,  Oregon. 
(Wickham.)  The  type  is  a  male  from  the  first  named  locality,  kindly 
given  me  by  Mr.  Waldo  Dodge  of  Melrose  Highlands,  Mass. 

This  species  is  remarkably  similar  in  nearly  all  respects  to  compar. 
The  entire  absence  of  elytral  clouds  (virtually  absent  in  some  speci- 
mens of  obscureplagiatus,  the  evident  disparity  in  elytral  punctures, 
and  the  generally  finer  and  less  dense  punctuation,  together  with  the 
locality  may  be  depended  on  to  distinguish  the  latter. 

C.  pedalis  Fall. 

A  little  larger  than  medialis  and  lutescens,  the  form  less  narrowed  and  hence 
more  obtusely  rounded  in  front.  Color  a  dingy  yellow,  the  head,  base  and 
diffuse  median  spot  on  the  prothorax,  and  almost  the  entire  disk  of  the  elytra 
infuscate;  beneath  black,  epipleurae  and  legs  dull  yellow.  Punctuation  above 
quite  dense,  a  little  less  so  basally,  the  irregular  series  of  coarser  punctures 
evident  on  the  elytra,  the  outer  discal  one,  however,  quite  inconspicuous. 

13 


Punctuation  beneath  coarse  and  dense,  the  epipleurae  much  more  densely  and 
completely  punctate  than  in  the  allied  species. 

Male :  front  and  middle  tarsi  very  little  wider  than  in  the  female,  the  claws 
of  the  front  tarsi  slender  and  similar;  front  femora  flattened  and  broadly  ob- 
liquely concave  distally  beneath,  middle  tibiae  sinuate  and  produced  inwardly 
at  apex,  hind  femora  without  the  usual  fine  strigosity,  coarsely  punctate  and 
obliquely  rugose  distally. 

Female :  legs  normal,  except  that  the  hind  femora  are  not  finely  strigose. 
Length  3.65  to  4.1  mm.;  width  2  to  2.15  mm. 

California.  Type  from  Pomona.  Known  to  me  also  from  Riverside, 
Naples,  San  Diego  (Blaisdell)  ;  Vine  Hill,  Contra  Costa  Co.  (Blaisdell.) 

The  peculiar  crural  sexual  modifications  make  this  one  of  the  most 
distinct  and  easily  recognized  species  of  the  genus  when  males  are  at 
hand.  Females  may  be  known  from  other  species  occurring  in  same 
region  by  the  more  obtuse  form  and  densely  punctate  epipleurae. 

C.  collatus  sp.  nov. 

Moderately  elongate  oval,  color  as  a  rule,  rather  clear  flavotestaccous  with  the 
usual  infuscations,  viz.;  the  outer  antennal  joints,  sides  of  the  front,  small 
discal  spot  and  basal  margin  of  the  prothorax  (more  or  less)  and  the  disk  of 
the  elytra;  the  broad  confluent  elytral  markings  distinctly  outlined  externally, 
the  suture  narrowly  pale,  at  least  toward  the  base.  Surface  shining  in  both 
sexes,  the  punctuation  typical  in  distribution  and  moderate  in  density.  Be- 
neath black,  polished,  with  the  usual  coarse  lateral  punctuation,  which,  how- 
ever, is  rather  finer  and  less  dense,  especially  on  the  coxal  plates,  than  in  such 
species  as  medialis,  suturalis  and  pedalis.  Front  and  middle  tarsi  of  male  mod- 
erately strongly  dilated,  the  second  joint  of  front  tarsi  not  wider  than  the 
first;  anterior  claw  of  front  tarsus  very  evidently  thickened  and  more  abruptly 
bent.  Length  3.75  to  4.2  mm.;  width  1.9  to  2.15  mm. 

The  type  is  a  male  taken  by  the  writer  at  Bear  Lake,  So.  California, 
8-3 1-' 1 6.  I  have  also  taken  the  species  at  Winslow,  Arizona,  and  have 
an  example  from  Cosnino  Canon,  Ariz.,  collected  by  Prof.  Wickham. 

This  species  is  closely  similar  in  nearly  every  way  to  patruelis,  but  is 
of  somewhat  larger  average  size,  with  a  little  less  strongly  dilated  male 
tarsi,  the  second  joint  not  wider  than  the  basal  one. 

C.  patruelis  Lee. 

Very  similar  to  collatus,  the  description  for  which  answers  well  enough  except 
as  follows :  Size  a  little  smaller,  females  of  two  forms — shining  and  opaque — 
the  elytral  cloud  often  more  diffuse  and  ill-defined,  though  frequently  quite  as 
sharply,  outlined  as  in  collatus.  Front  tarsi  of  male  more  strongly  dilated,  the 
second  joint  perceptibly  wider  than  the  basal  one.  Length  3.6  to  3.9  mm.;  width 
1.8  to  2  mm. 

Le  Conte's  type  was  from  "Fort  Laramie,  Nebraska."  Specimens 
at  hand  are  from  Cheyenne,  Wyoming  (Wickham),  and  Winnipeg, 
Stony  Mountain,  Miami,  and  Thornhill,  Manitoba,  all  collected  by  Mr. 

Wallis. 

14 


0.  infuscatus  Sharp. 

"Oblongo-ovalis,  fere  sine  pubescentia,  testaceous,  abdomine  pectoreque  nigris, 
elytris  signatures  magnis  coalescentibus  fuscis,  dense,  aequaliter  subtiliter  punc- 
tatis,  serie  punctorum  impressa  minus  distincta;  coxis  posterioribus  externe 
minus  crebre  punctatis.  Long.  5,  lat.  2%  mm." 

The  above  is  Sharp's  description  of  this  species,  which  is  unknown 
to  me  in  nature.  The  author  goes  on  to  remark  that  "This  species  is 
readily  distinguished  from  Hydroporus  nubilus  by  its  larger  size,  and 
more  finely  and  densely  punctured  elytra.  I  have  seen  only  two  in- 
dividuals, both  of  which  are  females,  one  is  shining,  the  other  dull ; 
the  front  tarsi  are  less  compressed  and  are  broader  than  in  the  corres- 
ponding sex  of  nubilus."  The  very  large  size,  together  with  the  lo- 
cality, will  probably  ensure  the  recognition  of  this  species  when  it  again 
turns  up.  Only  three  species  of  our  fauna — unguicularis,  impresso- 
punctatus  and  orcgonus — attain  a  length  of  5  mm. 

C.  nubilus  Lee. 

Elongate  oval,  yellowish  testaceous,  antennae  scarcely  at  all  infuscate  apically; 
head  pale,  with  or  without  a  small  transverse  fuscous  spot  contiguous  to  the 
thoracic  margin;  prothorax  with  small  discal  spot,  the  front  and  rear  margins 
scarcely  darker;  elytra  with  somewhat  diffuse  posteriorly  confluent  markings, 
consisting  of  two  subentire  vittae  near  the  suture,  exterior  to  which  are  an- 
terior and  posterior  spots  representing  two  more  abbreviated  and  interrupted 
vittae.  Elytral  punctuation  rather  close  and  subequal  throughout,  the  series 
of  coarser  punctures  nearly  or  quite  absent.  Beneath  black  moderately  punc- 
tate, epipleurae  more  closely  and  uniformly  so  than  in  patruelis.  Front  and 
middle  tarsi  of  male  quite  strongly  dilated,  the  second  joint  perceptibly  wider 
than  the  first  and  third ;  anterior  claw  of  front  tarsus  (G?)  a  little  shorter, 
thicker  and  more  strongly  bent  than  its  fellow.  Length  4  to  4.4  mm. ;  width 
2  to  2.2  mm. 

A  common  and  widely  diffused  species,  occurring  from  New  England 
to  Western  Texas.  Le  Conte's  type  was  from  Fort  Laramie  and  he 
speaks  of  its  occurrence  also  in  Illinois.  Specimens  seen  by  me  are 
from  Massachusetts  (Blanchard)  ;  N.  Y. ;  Jeannett,  Pa.,  Detroit,  Mich. ; 
McPherson  Kans. ;  Fedor,  Tex.  and  El  Paso,  Tex. 

The  elongate  form,  rather  large  size,  evenly  rather  finely  punctate 
elytra,  and  pale  head,  form  a  combination  of  characters  which  separate 
this  species  from  all  others  except  femoratus,  from  which  males  are 
easily  separable  by  the  tabular  characters.  In  the  series  of  nubilus 
before  me,  the  few  females  are  duller  and  finely  alutaceous ;  it  is,  how- 
ever, not  unlikely  that  shining  females  occur. 

C.  femoratus  Fall. 

This  species  is  so  exactly  similar  to  nubilus  as  to  be  quite  indistinguishable 
except  by  reference  to  the  femoral  and  tibial  structure  of  the  male.  In  this 
sex  all  the  femora  are  sinuate  beneath  near  the  knee,  the  margins  strongly 
laminate  at  apex.  The  tibiae  are  pedunculate  at  base,  then  suddenly  widened 


and  nearly  parallel,  instead  of  gradually  widening  from  base  to  apex  as  is 
usual.  Length  4.5  mm. ;  width  2.2  mm. 

The  type  is  a  male,  collected  at  Albuquerque  New  Mexico,  by  Prof. 
Wickham.  A  second  example,  taken  at  same  place  is  a  female  and  is 
probably  correctly  associated ;  it  is  slightly  larger,  and  noticeably  duller 
than  the  male. 

C.  fraternus  Lee. 

Rather  broadly  oval,  pale  flavo-testaceous  above,  black  beneath ;  males  shining, 
females  dull.  Antennae  scarcely  at  all  infuscate  at  tip.  Head  with  or  without 
a  small  occipital  spot;  prothorax  with  small  median  spot,  and  middle  of  base 
narrowly  darker,  both,  however,  sometimes  obsolete;  elytra  with  small  discal 
markings,  mostly  behind  the  middle.  Punctuation  of  elytra  rather  close  and 
even,  the  series  of  larger  punctures  scarcely  evident.  Beneath  coarsely  closely 
punctate.  Front  and  middle  tarsi  of  male  widely  dilated,  the  second  joint  per- 
ceptibly wider  than  the  first  and  third,  anterior  claw  of  front  tarsus  more 
strongly  curved  but  not  much  thicker  than  the  posterior  one.  Length  3.4  to 
3.95  mm.;  width  1.9  to  2.2  mm. 

California  ("New  River,  Colorado  Desert")  (type)  ;  Mokelumne  Hill 
(Blaisdell)  ;  Arizona  (Phoenix,  Palomas.) 

This  species  seems  quite  characteristic  of  the  arid  and  semi-arid 
regions  of  Arizona  and  contiguous  territory.  Its  occurrence  at  Moke- 
lumne Hill  on  the  western  side  of  the  California  Sierras  was  hardly 
to  be  expected.  The  species  is  most  likely  to  be  confounded  with  the 
paler  forms  of  medialis,  which  occur  in  the  same  region;  the  latter 
however  is  rather  narrower  in  form,  with  broader  elytral  clouds,  the 
tarsi  of  the  male  less  dilated  and  with  somewhat  differently  shaped 
anterior  claw,  the  females,  as  a  rule,  more  shining.  All  females  of 
fraternus  that  I  have  seen  are  alutaceous  and  dull. 

C.  tumidiventris  sp.  nov. 

Oval,  moderately  convex ;  head,  thorax  and  legs  rufotesiaceous,  elytra  pale 
flavotestaceous  to  dull  yellow,  body  beneath  black.  Head  black  or  fuscous  at 
sides  and  vertex ;  prothorax  with  basal  and  apical  margins  sometimes  narrowly 
infuscate,  a  small  fuscous  discal  spot,  rarely  lacking;  elytra  with  variable 
blackish  or  fuscous  markings,  which  in  the  type  consist  of  four  well  defined 
vittae  on  each,  the  two  inner  ones  entire,  the  two  outer  interrupted;  but  in  the 
greater  number  of  specimens  the  vittae  are  more  or  less  completely  fused 
posteriorly  or  even  throughout.  Antennae  with  the  outer  joints  lightly  in- 
fuscate at  their  apices.  Integuments  polished  in  the  male,  in  the  female  the 
surface  may  be  either  shining  like  the  male  or  finely  alutaceous  and  dull  both 
above  and  beneath.  Head  and  thorax  finely  sparsely  punctate,  elytra  with 
intermixed  fine  and  coarser  punctures,  the  sutural  and  two  discal  series  of 
coarser  punctures  distinct  as  far  as  the  middle.  Beneath  moderately  punctate ; 
last  ventral  obtusely  tumid  each  side,  the  tumidity  stronger  and  more  punctate 
in  the  male.  Front  and  middle  tarsi  of  male  rather  strongly  dilated,  the  second 
joint  widest,  but  less  conspicuously  so  than  in  the  three  preceding  species. 
Length  4.2  to  4.5  mm.;  width  2.15  to  2.4  mm. 

16 


Described  from  a  series  of  5d"s  and  4?'s,  two  of  the  latter  opaque.  The 
type  (c?)  bears  label  "Stony  Mountain,  Man.  I5-IV-I2,  J.  B.  Wallis." 
Other  localities  represented  are  Edmonton,  Alberta,  April  8,  1916 
(F.  S.  Carr)  ;  Wy. ;  Glenwood  Springs,  Colorado  (Fenyes)  ;  and 
Virgin  River,  Utah  (Roberts  Coll.) 

The  rather  large  size,  dual  elytral  punctuation  and  tumid  terminal 
ventral  segment  perfectly  define  this  species. 

C.  punctilineatus  sp.  nov. 

Oval,  pale  flavotestaceous  above,  black  beneath,  surface  (?)  both  above  and 
beneath  alutaceous  and  dull.  Head  black  each  side,  leaving  a  triangular  frontal 
area  pale :  thorax  with  a  small  discal  dot  and  the  extreme  edge  of  the  basal 
margin  at  middle,  piceous ;  elytra  each  with  four  narrow  vittae  and  the 
suture,  black ;  outer  vittae  interrupted  at  middle.  Punctuation  above  very  fine, 
with  coarser  punctures  along  the  suture  basally,  and  elsewhere  for  the  most  part 
confined  to  the  black  vittae,  especially  apically.  Beneath  moderately  punctate, 
most  strongly  and  closely  on  the  coxal  plates.  Length  3.6  mm.;  width  1.95  mm. 

Described  from  a  single  female  specimen  taken  by  Mr.  J.  B.  Wallis 
at  Stony  Mountain,  Manitoba,  April  13,  1912. 

The  front  tarsi  of  the  female  are  moderately  dilated  and  it  is  prob- 
able that  they  are  quite  widely  so  in  the  male.  This  species  is  most 
closely  allied  to  tumidwentris,  but  the  size  is  notably  smaller,  the 
punctuation  very  much  finer,  the  coarser  punctures  of  the  elytra  fewer 
and  differently  disposed,  and  the  last  ventral  without  trace  of  the 
apical  tumidities. 

C.  masculinus  Cr. 

Elongate,  slightly  obovate,  elytra  pale  flavotestaceous,  head  and  thorax  a  little 
more  rufous,  beneath  black.  Surface  shining  in  the  male,  finely  alutaceous  and 
dull  in  the  female.  Punctuation  of  head  and  thorax  fine,  not  dense,  very  slightly 
coarser  along  the  base  of  the  latter;  elytra  finely,  quite  densely  and  very 
evenly  punctate,  the  series  of  coarser  punctures  entirely  lacking.  Head  entirely 
pale,  clypeus  finely  margined.  Prothorax  a  little  narrower  at  base  than  the 
contiguous  base  of  the  elytra,  sides  feebly  arcuate  or  nearly  straight,  not  very 
strongly  convergent  in  front  and  not  continuous  with  the  sides  of  the  elytra ; 
base  very  narrowly  dusky  at  middle,  otherwise  entirely  pale.  Elytra  each 
with  four  narrow  vittae  and  the  suture,  black ;  vittae  i  and  3  abbreviated  at  base, 
4  abbreviated  apically,  3  and  4  with  a  tendency  toward  interruption  at  middle. 
Body  beneath  finely  alutaceous,  sides  of  metasternum,  sides  of  abdomen  at  base, 
and  last  ventral  segment  more  coarsely  punctate,  elsewhere  sparsely  finely  punc- 
tate. Anterior  tarsi  of  male  rather  strongly  dilated,  the  claws  very  long, 
broadly  laminiform,  dilated  at  middle,  the  anterior  one  a  little  shorter  and 
less  acute  at  tip.  Length  4.5  mm.;  width  2.3  mm. 

Crotch's  type  was  described  from  Lake  Lebache,  the  precise  location 
of  which  is  unknown  to  me  but  presumably  in  British  Columbia  or 
Alberta.  In  my  own  collection  this  quite  remarkable  species  is  rep- 
resented by  a  single  pair,  the  male  from  Dakota  and  female  from  Win- 
nipeg, Man.  (Wallis.) 

17 


In  addition  to  the  several  characters  given  in  the  table,  it  may  be 
added  that  in  this  and  the  following  species,  the  spines  along  the  inner 
edge  of  the  basal  joint  of  the  hind  tarsus  are  shorter  and  more  close 
set  than  in  any  other  species  known  to  me. 

C.  fastidiosus  sp.  nov. 

Very  closely  allied  to  masculinus  and  possibly  to  be  united  with  it  as  a  local 
variety  when  further  specimens  including  males  are  known.  The  form  is 
a  little  less  distinctly  obovate,  the  prothorax  larger,  sides  longer,  just  perceptibly 
sinuate,  and  continuous  with  the  sides  of  the  elytra,  feebly  subexplanate,  es- 
pecially toward  the  anterior  angles;  not  in  the  least  so  in  masculinus.  Length 
4.7  mm. ;  width  2.4  mm. 

A  single  female  specimen  collected  at  Owen's  Lake,  California,  V-22- 
17  by  G.  R.  Pilate. 

C,  semivittatus  sp.  nov. 

Moderately  elongate,  slightly  obovate,  pale  flavotestaceous,  the  head  and  thorax 
of  slightly  brighter  tint  and  entirely  immaculate;  elytra  each  narrowly  quadri 
vittate  with  fuscous  posteriorly,  the  two  inner  vittae  on  each  extending  to  a 
point  a  little  in  advance  of  the  middle,  the  two  outer  ones  represented  only  by 
one  or  two  short  fragments,  the  suture  also  narrowly  dark,  surface  (?)  very 
finely  alutaceous  and  dull  both  above  and  beneath.  Antennae  with  the  outer 
joints  partly  infuscate.  Prothorax  small,  strongly  transverse,  sides  nearly 
straight,  only  slightly  divergent  posteriorly  and  not  continuous  in  outline  with 
the  elytra.  Head  and  thorax  very  finely,  not  closely  punctate,  elytra  densely 
evenly  so,  almost  without  trace  of  coarser  punctures.  Length  4.5  mm.;  width 
2.2  mm. 

The  unique  female  type  was  taken  by  the  writer  at  Seligman, 
Arizona,  July  3,  1906. 

The  four  anterior  tarsi  are  narrow,  the  first  three  joints  elongate 
parallel  and  of  equal  width.  It  is  quite  possible  that  further  specimens 
will  show  that  the  elytral  vittae  are  not  constant  in  their  degree  of 
development.  The  somewhat  obovate  form  and  small  thorax  with 
sides  discontinuous  in  outline  with  the  elytra,  are  quite  suggestive  of 
masculinus,  but  the  structure  of  the  clypeus,  prosternal  process  and 
basal  joint  of  hind  tarsus  are  normal  in  the  present  species. 

C.  hudsonicus  sp.  nov. 

Moderately  elongate  oval  or  feebly  obovate,  dull  yellowish  testaceous,  head 
more  or  less  infuscate  at  sides,  thorax  witth  small  discal  spot,  elytra  each  with 
four  vittae  and  the  suture  piceous,  body  beneath  black.  Upper  surface  of  male 
finely  reticulato-alutaceous  and  moderately  shining,  in  the  female  more  dis- 
tinctly alutaceous  and  dull.  Elytr-a  evenly,  rather  closely  punctate,  coarser 
punctures  scarcely  evident;  head  and  thorax  more  finely  and  sparsely  punctate 
as  usual.  Beneath  alutaceous  and  rather  sparsely,  not  very  coarsely  punctate. 
Front  and  middle  tarsi  of  male  broadly  dilated,  the  second  joint  widest,  the 
claws  more  elongate  than  usual,  the  anterior  one  curved  and  a  little  shorter 
than  the  posterior,  which  is  nearly  straight.  In  the  female  the  claws  of  the 
front  tarsi  are  much  shorter  than  in  the  male,  and  equal.  Length  3.8  to  4 
mm.;  width  1.85  to  1.95  mm. 

IS 


Described  from  a  single  pair  (the  female  smaller)  taken  at  Ungava 
Bay,  H.  B.^T.,  by  L.  M.  Turner. 

A  quite  distinct  species,  though  evidently  allied  to  unguicularis.  The 
latter  is  much  larger,  less  elongate,  and  with  different  front  claws  in 
the  male. 

C.  unguicularis  Cr. 

Form  oval,  not  or  scarcely  twice  as  long  as  wide,  fulvotestaceous  above,  black 
beneath.  Male  shining,  female  dull.  Head  at  the  extreme  base,  a  small  discal 
spot  and  the  middle  of  the  base  of  the  prothorax,  and  the  suture  and  four 
vittae  on  each  elytron  blackish  or  fuscous.  The  elytral  vittae  are  rather  wide, 
the  outermost  one  shortest,  interrupted  at  middle  and  often  more  or  less 
confluent  witth  the  next  inner  one.  Head  and  thorax  not  very  closely  punc- 
tulate,  the  punctures  a  little  closer  along  the  thoracic  base ;  elytra  densely,  rather 
finely  punctate,  without  series  of  coarser  punctures.  Beneath  alutaceous  in  both 
sexes,  punctuation  moderate  in  coarseness,  typical  in  distribution.  Front  tarsi 
of  male  broadly  dilated,  the  second  joint,  not  or  only  just  perceptibly  wider 
than  the  first  both  claws  stout,  acuminate,  the  anterior  one  but  little  more  than 
half  as  long  as  the  other.  Length  5  to  5.5  mm. ;  width  2.6  to  2.8  mm. 

Crotch's  type  was  from  British  Columbia.  Specimens  at  hand  are 
from  Winnipeg,  Le  Pas,  and  Mile  214  H.  B.  Ry.,  Manitoba;  Overland 
Lake,  Boulder  Co.,  Colorado  (Rohwer.) 

This  is  a  very  distinct  and  easily  recognized  species  when  due  at- 
tention is  paid  to  its  characters.  It  is  our  largest  species  with  the 
exception  of  impressopunctatus  and  or  eg  onus.  Certain  examples  of 
tumidiv  entris  approach  the  smaller  specimens  of  unguicularis  and  look 
quite  similar  but  the  resemblance  is  only  superficial. 

C.  oregonus  sp.  nov. 

Elongate  oval,  dull  yellowish  testaceous,  shining  (d1),  elytra  each  with  four 
rather  wide  fuscous  stripes  showing  a  tendency  to  confluence,  the  suture  also 
very  narrowly  dark.  Head  and  thorax  pale,  almost  throughout,  a  small  discal 
fuscous  spot  on  the  latter.  Punctuation  of  elytra  moderately  close,  rather  fine, 
without  distinct  series  of  coarser  punctures;  head  and  thorax  more  finely  and 
sparsely  punctate  as  usual.  Body  beneath  black,  not  alutaceous,  moderately 
coarsely  punctate.  Front  tarsi  of  male  broadly  dilated,  the  second  joint  scarcely 
visibly  wider  than  the  first,  claws  short,  stout,  evenly  curved  and  mutually 
equal.  Length  5.8  mm.;  width  2.85  mm. 

Corvallis,  Oregon.  A  single  male  specimen  submitted  by  Mr. 
Sherman,  who  retains  the  type. 

Slightly  larger  and  more  elongate,  but  very  similar  to  unguicularis 
superficially.  The  size,  vittate  elytra,  shining  non-alutaceous  lower 
surface,  and  short,  stout,  equal  claws  of  male  front  tarsi  are  sufficient 
for  its  easy  recognition. 

C.  impressopunctatus  Schall. 

Oblong  oval,  above  yellowish  testaceous,  to  dark  ferruginous,  head  at  sides 
and  behind,  and  base  of  thorax  more  or  less  infuscate;  elytra  either  more  or 

19 


less  evidently  vittate  or  with  the  vittae  suffused  into  a  broad  dorsal  cloud,  or 
with  the  entire  surface  brownish  piceous;  beneath  black.  Head  finely  punctate; 
prothorax  unevenly  punctured,  finely  so  in  front,  with  numerous  coarser 
punctures  in  the  basal  region;  elytra  in  the  male  with  fine  and  much  coarser 
punctures  conspicuously  intermixed  with  the  usual  discal  lines  of  coarse 
punctures  regular  and  impressed  in  basal  half,  becoming  obsolete  at  or  behind 
the  middle.  In  the  female  the  elytra  are  generally  more  densely  punctate  and 
dull,  the  intermixed  fine  punctures  less  evident  and  the  impressed  discal  lines 
feeble  or  wanting;  some  females,  however,  have  the  elytra  shining  and 
punctate  as  in  the  male.  Beneath  moderately  coarsely  punctate,  alutaceous, 
opaque  in  the  dull  females,  more  shining  in  the  males.  Tarsi  rather  broad  in 
both  sexes,  but  evidently  wider  in  the  male,  the  basal  joint  slightly  narrower 
than  the  second,  the  anterior  claw  of  the  front  tarsus  (c?)  thicker  and  more 
strongly  curved,  apparently  shorter  than  its  fellow  when  viewed  from  the 
front.  Length  4.5  to  5.4  mm.;  width  2.3  to  2.7  mm. 

A  very  widely  dispersed  species,  occurring  from  New  England  to 
Alaska  in  this  country,  as  well  as  in  Siberia,  Asia  Minor  and  Northern 
Europe. 

There  is  a  great  deal  of  variation  in  form,  color,  markings  and 
sculpture  as  might  be  expected  in  a  species  of  circum-terrestrial  dis- 
persion. 


20 


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